The world’s most popular cat, nicknamed Grumpy Cat, has died aged seven, her owners say.
A statement says she died on Tuesday following complications from a recent urinary tract infection.
The cat from Arizona had “helped millions of people smile”.
Grumpy’s real name was Tardar Sauce.
The cat became went viral in 2012 after photographs of her sour expression emerged online. Her image quickly spread as a meme.
Grumpy Cat travelled the world making television appearances and in 2014 even starred in her own Christmas film.
Madame Tussauds in San Francisco unveiled a waxwork of her in 2015.
Her Instagram account has more than two million followers.